| Collector | Holm-Nielsen, L. B. |
| Number | 5385 |
| Collected with | Jeppesen, S. & Lojtnant, B. & Ollgaard, B. |
| Country | Ecuador |
| Subpart | CARCHI |
| County | |
| Locality | Páramo El ángel, in the pass on road El ángel-Tulcán. Very humid Espeletia-paramo. |
| Coordinates | 77° 54' N 0° 41' W |
| Minimum elevation (m) | 3750 |
| Maximum elevation (m) | 3850 |
| Start date | 1973/05/15 |
| End date | 1973/05/15 |
| Family | Marchantiophyta |
| Identification | Frullania atrata Nees |
| Determined by | W. C. Steere |
| Identification qualifier | |
| Field notes | Epiphytic in cloud forest. |
This record